DUPLICATIONS # Doubler: see Dp(1;1)BSRMG # Dp(1)3C1: see Dp(1;1)wrG # Dp(1;1)3C3 cytology: Dp(1;1)3C2-3;3C3-5; tandem duplication of right-most sites of w (we and wsp). synonym: Dp3C3; Dp(w+R); Dp(we;wsp). references: Green, 1963, Z. Vererbungsl. 94: 200-14. 1967, Biol. Zentralbl. 86: 211-20. Arcos-Teran, 1972, Chromosoma 37: 233-96. genetics: Homozygotes wild type in eye color. Males and females with z and the duplication are z. # Dp(1;1;1)3C3 cytology: Triplication of right-most sites of w inferred (Green, 1967). No thickening of 3C3 visible in polytene chro- mosomes (Lefevre). origin: From homozygous Dp(1;1)3C3 females. synonym: Tr(w+R); Dp(1;1)3C3 (triplication). references: Green, 1967, Biol. Zentralbl. 86: 211-20. Arcos-Teran and Beerman, 1968, Chromosoma 25: 377-91. Arcos-Teran, 1972, Chromosoma 37: 233-96. genetics: Homozygotes wild type in eye color. Males and females with z and the duplication are z. # Dp(1;1)4E;6F cytology: Dp(1;1)4E2-F1;6F1-2. Reversed repeat. new order: 1A - 6F|6F - 4E|6F - 20. origin: Derived from Uc unstable X. references: Lim, 1979, Genetics 93: 681-701. genetics: Unstable. #*Dp(1;1)100: Duplication (1;1) 100 origin: Spontaneous product of exchange between Dp(1;f)100 and proximal heterochromatin of C(1)RM. #*Dp(1;1)105 cytology: Metaphase X chromosome has one arm of normal length and one about 40% normal length. new order: 1 - 20F.|6 - 1. origin: X-ray-induced deletion of most of X euchromatin was recovered as a C(1)RM/Dp(1;f)105 female, which by detachment produced Dp(1;1)105 in the succeeding generation. discoverer: Dobzhansky, 1930. references: 1932, Biol. Zentralbl. 52: 493-509. genetics: Contains wild-type alleles of y through dx and also probably bb. # Dp(1;1)112 origin: Spontaneous product of exchange between Dp(1;f)112 and proximal heterochromatin of an attached-X. #*Dp(1;1)138 origin: X-ray-induced deletion of most of X euchromatin that was recovered as a C(1)RM/Dp(1;f)138 female, which by detach- ment produced Dp(1;1)138 in the subsequent generations. discoverer: Dobzhansky, 1930. references: 1935, Z. Indukt. Abstamm. Vererbungsl. 68: 134- 62. genetics: Extends from locus of r to base of X; carries B. Female nearly wild type, but male has low viability and is sterile. #*Dp(1;1)258-46 cytology: Dp(1;1)2B4-7;3A4-6; reversed repeat (Sutton). new order: 1 - 2B4|3A4 - 2B7|2B7 - 20 or 1 - 3A4|3A4 - 2B7|3A6 - 20. origin: X ray induced. discoverer: Demerec, 38l. genetics: Originally appeared as w but reverted to w+. No effect on pn-gt reported. # Dp(1;1)Ax28a: Duplication (1;1) Abruptex cytology: Single-band duplication believed to involve 3C7 by Morgan, Schultz, and Curry (1941); identity of extra band not confirmed by Lefevre (1953, Genetics 38: 345-59). new order: 1A - 3C7|3C7 - 20. origin: X ray induced. references: Morgan, Schultz, and Curry, 1941, Year Book - Car- negie Inst. Washington Welshons, 1971, Genetics 68: 259-68. genetics: Ax phenotype. # Dp(1;1)B: Duplication (1;1) Bar cytology: Dp(1;1)15F9-16A1;16A7-B; a tandem duplication (Bridges, 1936, Science 83: 210-11; Muller, Prokofyeva- Belgovskaya, and Kossikov, 1936, Dokl. Acad. Nauk SSSR 1: 87-88). new order: 1 - 16A7|16A1 - 20. origin: Spontaneous. discoverer: Tice, 13b. references: 1914, Biol. Bull. 26: 221-30. genetics: Position effect for B, apparently resulting from jux- taposition of 16A1 with 16A7, which may undergo mutation to less extreme forms (e.g., Bi). Produces normal and tripli- cated [Dp(1;1;1)BB] products by unequal crossing over. #*Dp(1;1)B263-28 cytology: Dp(1;1)15F9-16A1;16A3-4;16A6-7;16A7-B1. new order: 1 - 16A3|16A7|16A1 - 20. origin: X-ray-induced deletional derivative of Dp(1;1)BiBi = Dp(1;1)15F9-16A1;16A7-B1. discoverer: Demerec, 34b. references: Sutton, 1943, Genetics 28: 97-107. #*Dp(1;1)B263-48: see Tp(1;1)B263-48 # Dp(1;1)BS-H: Duplication (1;1) Bar Stone of Hinton new order: 1A1-1B3|20F-3C2|20F|1A4-3C1|20F-16A1|102F; position of cen- tromere indeterminate; adjacent to either 3C1 or 3C2. origin: Generated by double exchange in a R(1)2, In(1)wvC/In(1)sc4Lsc8R/XP4D female, one exchange occurring between the BS element and R(1)2 and the other between In(1)sc4Lsc8R and R(1)2. references: Hinton, 1957, Genetics 42: 55-65. phenotype: These linear derivatives of the wvC ring do not undergo the somatic loss characteristic of the unstable ring; they do, however, display indications of variable levels of instability, presumably related to the degree of instability of the generating ring. This is recognized by reduced recovery and elevated primary nondisjunction in Dp(1;1)BS-H/In(1)dl49 females. Exchange between the BS dupli- cation and the long arm of the same chromatid regenerates sin- gle rings whose stabilities reflect those of the parental linear chromosome. Exchange between the BS duplication and a normal homologue generates tandem-metacentric compound chromo- somes [C(1)TM-H]. # Dp(1;1)BSRAG: Duplication (1;1) Bar of Stone Reversed Acrocentrigenic cytology: Dp(1;1)15F9-16A1;20. new order: .20|1A - 20|20 - 16A1|102F. origin: Spontaneous recombinant between the distal X of a C(1)RA and XP4D from T(1;4)BS. discoverer: Lindsley and Sandler. references: 1963. Methodology in Basic Genetics (W. J. Burdette, ed.). Holden-Day, Inc., San Francisco, pp. 390-403. genetics: Generates reversed acrocentric compound X chromosomes in Dp(1;1)BSRAG/+ female, usually by a double exchange in which one exchange occurs between the duplicated segment of one strand and the homologous region of its sister and the other between the duplication-bearing X and its normal homo- log. Rate of C(1)RA generation about 6 x 10-4. # Dp(1;1)BSRMG: Duplication (1;1) Bar of Stone Reversed Metacentrigenic cytology: Dp(1;1)15F9-16A1;20. new order: 1 - 20.20 - 16A1|102F. origin: Spontaneous recombinant between C(1)RM and the XP4D element of T(1;4)BS. discoverer: Muller. synonym: Doubler. references: 1936, DIS 6: 8. Lindsley and Sandler, 1963, Methodology in Basic Genetics (W. J. Burdette, ed.). Holden-Day, Inc., San Francisco, pp. 390-403. genetics: Generates reversed metacentric compound X chromosomes in Dp(1;1)BSRMG/+ female by crossing over between the dupli- cated segment and either the X to which it is attached or the homologous X at a rate of about 2.5 x 10-4. # Dp(1;1)BSTAG: Duplication (1;1) Bar of Stone Tandem Acrocentrigenic cytology: Dp(1;1)15F9-16A1;20. new order: .20 - 1A|20 - 16A1|102F. origin: X-ray-induced recombinant between the distal hetero- chromatin of an X chromosome with a terminal heterochromatic segment derived from y+Y and the proximal heterochromatin of the XP4D element of T(1;4)BS. discoverer: Lindsley and Sandler. references: 1963, Methodology in Basic Genetics (W. J. Burdette, ed.). Holden-Day, Inc., San Francisco, pp. 390-403. genetics: Ineffective in generating tandem acrocentric compound X chromosomes. # Dp(1;1)BSTMG: Duplication (1;1) Bar of Stone Tandem Metacentrigenic cytology: Dp(1;1)15F9-16A1;20 added as a second arm to In(1)sc8LENR. new order: 1A - B2|20B - 1A|20.20 - 16A1|102F. origin: Spontaneous recombinant between the X in normal sequence of a C(1)TM and the XP4D element of T(1;4)BS. discoverer: Lindsley and Sandler. references: 1963, Methodology in Basic Genetics (W. J. Burdette, ed.). Holden-Day, Inc., San Francisco, pp. 390-403. genetics: Generates tandem metacentric compound X chromosomes in Dp(1;1)BSTMG/+ female by recombination between the duplica- tion and the base of a homolog in normal sequence, at a rate of about 20 x 10-4. # Dp(1;1)BSTRG: Duplication (1;1) Bar of Stone Tandem Ring-genic See C(1)TMBS subsection in compound chromosomes. # Dp(1;1)bbDfLClR cytology: Dp(1;1)4A5-B1;4D2-3 + Dp(1;1)17A6-B1;20B-C. origin: Associated with In(1)bbDfLClR. references: CP627. # Dp(1;1)bbDfLy4R cytology: Dp(1;1)1A8-B1;4D2-3 + Dp(1;1)18A3-4;20B-C. origin: Associated with In(1)bbDfLy4R. references: CP627. # Dp(1;1;1)BB: Duplication (1;1;1) Bar + Bar cytology: Dp(1;1)15F9-16A1;16A7-B1; a tandem triplication [Bridges, 1936, Science 83: 210-11 (fig.)]. new order: 1 - 16A7|16A1 - 16A7|16A1 - 20. origin: Spontaneous through unequal crossing over in B/B female. discoverer: Zeleny. references: 1920, J. Exp. Zool. 30: 292-324. Sturtevant, 1925, Genetics 10: 117-47. genetics: Either or both B regions may carry a less extreme derivative of B (i.e., BiB, BBi, or BiBi). Number of dupli- cated segments may be either increased or decreased by unequal crossing over. #*Dp(1;1)Bt: Duplication (1;1) Branchlet cytology: Dp(1;1)3B2-C1;6F6-7; tandem repeat (Darby). new order: 1 - 6F4|3C1 - 20. origin: Induced by 32P. discoverer: Bateman, 1950. references: 1950, DIS 24: 54. 1951, DIS 25: 77. # Dp(1;1)Bxr: Duplication (1;1) Beadex-recessive cytology: Dp(1;1)17A;17E-F (E. B. Lewis). origin: Spontaneous. discoverer: Ives, 35k. references: 1937, DIS 7: 6. Green, 1952, Proc. Nat. Acad. Sci. USA 38: 949-53. 1953, Genetics 38: 91-105. 1953, Z. Indukt. Abstamm. Vererbungsl. 85: 435-49. genetics: Duplicated for os+, Bx+, and fu+. Does not yield unequal crossovers as does Bxr49k. # Dp(1;1)Bxr49k cytology: Dp(1;1)17A;17C (E. B. Lewis). origin: Spontaneous. discoverer: Mossige, 49k22. references: 1950, DIS 24: 61. Green, 1953, Z. Indukt. Abstamm. Vererbungsl. 85: 435-49. Green, 1968, Mol. Gen. Genet. 103: 209-17. genetics: Duplicated for Bx+ but not for os+ or fu+. Unequal crossing over yields wild types and triplications. Quadrupli- cations have also been produced. # Dp(1;1)ClLy4R cytology: Dp(1;1)1A8-B1;4A5-B1+Df(1)17A6-B1;18A3-4.. new order: 1 - 4A|17A6-1B1|18A4 - 20. origin: Associated with In(1)ClLy4R. references: CP627. # Dp(1;1)Co: Duplication (1;1) Confluens cytology: Dp(1;1)3C4-5;3D6-E1; tandem duplication (Schultz, 1941, DIS 14: 54-55). new order: 1 - 3D6|3C5 - 20. origin: Recovered among progeny of cold-treated fly. discoverer: Gottschewski, 34c. references: 1935, DIS 4: 7, 14, 16. 1937, DIS 8: 12. 1937, Z. Indukt. Abstamm. Vererbungsl. 73: 131-42. genetics: The Co phenotype arises from a duplication of the Notch locus (salivary band 3C7). # Dp(1;1)ct78 cytology: Dp(1;1)6F1-2;7C1-2. Reversed repeat. new order: 1 - 6F|7C - 6F|6F - 20. origin: Derived from Uc unstable X. references: Lim, 1979, Genetics 93: 681-701. genetics: Unstable. # Dp(1;1)G: Duplication (1;1) Gelbart references: Gelbart, 1971, Ph.D. Thesis. duplication cytology _________________________________ Dp(1;1)GJ4 1B6-7;3C1-2;3C2-3 ( Dp(1;1)GJ5 3C2;3C3 + In(1)1B;10A Dp(1;1)GM9 3B;3F Dp(1;1)GM10 3A2-3;3C3 ( new order: 1 - 1B6|3C2 - 1B7|3C2 - 20 (tentative). # Dp(1;1)Gr: Duplication (1;1) Green cytology: Dp(1;1)3A2-3;8B4-C1; tandem repeat. More than 1/4 the length of euchromatic X at metaphase. origin: Spontaneous. references: Kalisch, 1973, Chromosoma 41: 237-42. 1975, Theoret. Appl. Genet. 46: 169-80. 1976, Genet. Res. 26: 275-82. 1980, DIS 55: 206-07. Charton-Struck and Kalisch, 1981, DIS 56: 28-29. genetics: Homo- and hemizygous lethal. Heterozygous females viable; show reduction in crossing over within and adjacent to the duplication and increase in crossing over in distal parts of 2 and 3. Duplication chromosome marked by y2 sc distally and (w- spl sn3)(wc sn3) within the duplication. # Dp(1;1)hdp-b: Duplication (1;1) heldup-b origin: Induced in progeny of dysgenic flies. references: Engles and Preston, 1984, Genetics 107: 657-78. genetics: Mutant for hdp-b as result of break at 17C, site of P factor in strain ~~2. chrom. duplication cytology design. new order ________________________________________________________________________ Dp(1;1)hdp12 17C;17C Dp(1;1)hdp-b1 ( B395.2 1A - 17C|dp|17C - 20. Dp(1;1)hdp-b2 12F;13E;17C F14f 1A - 17C|12F - 13E|17C - 20. Dp(1;1)hdp-b3 5A;7A;17C;19F L40 1A - 7A|19F - 17C|5A - 17C| 19F - 20 Dp(1;1)hdp-b4 | N71 1A - 17C|dp|17C - 20. Dp(1;1)hdp-b5 ( N86 1A - 17C|dp|17C - 20. Dp(1;1)hdp-b6 4F;5E;17C N224 1A - 5E|17C - 4F|17C - 20. Dp(1;1)hdp-b7 4F;5E;17C N254 1A - 17C|5E - 4F|17C - 20. ( Dp for one of more bands. | Dp for several bands. # Dp(1;1)hdp-brv: Duplication (1;1) heldup-b reverted cytology: Dp(1;1)5E;7A; reversed repeat. new order: 1A - 5E|7A - 5E|5E - 20. synonym: Dp(1;1)019. origin: Induced in progeny of hdp-b fly (P factors in rear- ranged chromosome at 5E and 17C). references: Engels and Preston, 1984, Genetics 107: 657-78. genetics: Reverted for hdp-b. # Dp(1;1)IMNB-8 cytology: Inversion of segment overlapping the tandem duplica- tion Dp(1;1)MNB-8. new order: 1A - 16F1|16F - 16A1|16A7 - 16A1|17E1 - 17A2|16F - 20. origin: Spontaneous derivative of Dp(1;1)MNB-8. references: Holmquist, 1972, Chromosoma 36: 413-52. genetics: Has a B os+ phenotype and a (B+ B)(os+ os) genotype. # Dp(1;1)jnR1-A cytology: Small duplication in 6F-7A region. references: Mainz, Salz, Cline, and Schedl, 1985, Cell 43: 521-29. genetics: Duplicated for Sxl+ and l(1)7Aa. # Dp(1;1)jnR1-B cytology: Small duplication in 6F-7A region. references: Mainz, Salz, Cline, and Schedl, 1985, Cell 43: 521-29. genetics: Duplicated for l(1)7Aa but not Sxl+. # Dp(1;1)L-BS: Duplication (1;1) Lindsley origin: Replacement of y+ in Dp(1;1)L-y+ with BS from BSYy+ resulting in the production of a BS(het) y cv v f bb- chromo- some. genetics: Carries BS distal to and y cv v f bb- proximal to the heterochromatin. # Dp(1;1)L-y31d origin: Replacement of y+ in Dp(1;1)L-y+ with y31d from FM6 resulting in production of a y31d (het) y cv v f bb- chromo- some. genetics: Carries y31d distal to and y cv v f bb- proximal to the distal heterochromatin. # Dp(1;1)L-y+ origin: X-ray-induced recombination in a BSYy+/FM6/YSX, Df(1)bb3a, y cv v f female resulting in a YSX, Df(1)bb3a, y+ KS y cv v f bb- chromosome. references: Lindsley and Sandler, 1963, Methodology in Basic Genetics (Burdette, ed.). Holden-Day, Inc., San Francisco, pp. 390-403. genetics: Carries y+ distal to and y cv v f bb- proximal to the distal heterochromatin. # Dp(1;1)lz-1: Duplication (1;1) lozenge cytology: Dp(1;1)8D;8F; tandem duplication. origin: X ray induced in lz50e+/+lzy4 female. references: Bender, 1967, Genetics 55: 249-54. Peeples, Geisler, Whitcraft, and Oliver, 1969, Genetics 62: 161-70. genetics: Partial complementation for lz. Hemizygous males have slightly rough eyes, thin or missing posterior postalar bristles, and good fertility; homozygous females have rougher eyes, thin or missing posterior postalar bristles, and markedly reduced fertility although secondary reproductive structures are normal. Tarsal claws are normal in both sexes. # Dp(1;1)lz-2 cytology: Dp(1;1)8D1+; small tandem duplication. origin: X ray induced in lz50e+/+lzy4 female. references: Bender, 1967, Genetics 55: 249-54. Peeples, Geisler, Whitcraft, and Oliver, 1969, Genetics 62: 161-70. De la Concha, Dietrich, Weigel, and Campos-Ortega, 1988, Genetics 118: 499-508. genetics: Partial complementation for lz. Duplication may include t+. Dp(1;1)lz-2 males have less rough eyes than Dp(1;1)lz-1 males, normal posterior postalar bristles, and good fertility; homozygous Dp(1;1)lz-2 females have moderately rough eyes, thin or missing posterior postalar bristles, and good fertility; tarsal claws normal in both sexes. # Dp(1;1)MNB-8 cytology: Dp(1;1)15F9-16A1;16A7-B1;17E2-F1 + Df(1)16 E- F1;17A2-3; tandem repeat with three copies of 16A1-7. new order: 1A - 16E|17A3 - 17E1|16A1 - 16A7|16A1 - 20. origin: Spontaneous derivative of Dp(1;1)NB-8. synonym: MLD, modified long duplication. references: Gabay and Laughnan, 1970, Genetics 65: 249-65. Laughnan, Gabay, and Montgomery, 1971, DIS 47: 64. Holmquist, 1972, Chromosoma 36: 413-52. genetics: Has a B os+ phenotype and a (B+ os+) (B os) genotype. Intrachromosomal recombination in some lines results in excep- tional male offspring (Peterson and Laughnan, 1964, Genetics 50: 275-76; Gabay and Laughnan, 1970; Laughnan et al., 1971). # Dp(1;1)NB-8 cytology: Dp(1;1)15F9-16A1;16A7-B1;17E2-F1; tandem repeat with three copies of 16A1-7. new order: 1A - 17E1|16A1 - 16A7|16A1 - 20. origin: Derived from Dp(1;1)NBB-8 by exchange with wild type X (crossover in distal 16A1-17E2 segment of the duplication). synonym: SLD, standard long duplication. references: Gabay and Laughnan, 1970, Genetics 65: 249-65. genetics: Has a B os+ phenotype and a (B+ os+) (B os) geno- type. #*Dp(1;1)NBB-8 cytology: Dp(1;1)15F9-16A1;17E2-F1 superimposed on Dp(1;1)15F9-16A1;16A7-B1; four copies of 16A1-7. new order: 1 - 16A7|16A1 - 17E2|16A1-16A7|16A1 - 20. origin: Spontaneous in a B/f B os car female. discoverer: Peterson and Laughnan. synonym: OLD, original long duplication. references: 1963, Proc. Nat. Acad. Sci. USA 50: 126-33. Gabay and Laughnan, 1970, Genetics 65: 249-65. genetics: Male viability reduced. Has a BB phenotype. Geneti- cally it is (B os+) (B os). # Dp(1;1)O19 cytology: Dp(1;1)5E;7A. new order: 1A - 5E|7A - 5E|5E - 20. origin: Induced by hybrid dysgenesis. references: Engles and Preston, 1984, Genetics 107: 657-78. # Dp(1;1)pn2: Duplication (1;1) prune synonym: TE100. genetics: Duplicated for Pgd. # Dp(1;1)S93: see Tp(1;1)S93 # Dp(1;1)sc4Lsc8R cytology: Dp(1;1)1B2-3;1B3-4. origin: Associated with In(1)sc4Lsc8R. # Dp(1;1)sc4Ly4R cytology: Dp(1;1)1A8-B1;1B3-4 + Dp(1;1)18A3-4;h26. origin: Associated with In(1)sc4Ly4R. # Dp(1;1)sc8Lsc4R cytology: Dp(1;1)h26;h32. origin: Associated with In(1)sc8Lsc4R. # Dp(1;1)sc8LscL8R cytology: Dp(1;1)h30-31;h32. origin: Associated with In(1)sc8LscL8R. # Dp(1;1)sc8Ly4R cytology: Dp(1;1)1A8-B1;1B2-3 + Dp(1;1)18A3-4;h32. origin: Associated with In(1)sc8Ly4R. # Dp(1;1)sc260-25 origin: Aneuploid recombinant from In(1LR)sc260-25. # Dp(1;1)scL8Lsc4R cytology: Dp(1;1)h26;h30-31. origin: Associated with In(1)scL8Lsc4R. # Dp(1;1)scL8Lsc8R cytology: Dp(1;1)1B2-3;1B3-4. origin: Associated with In(1)scL8Lsc8R. # Dp(1;1)scS1Lsc4R (L. Robbins) cytology: Dp(1;1)20F;1B3-4. NO and portions of surrounding heterochromatin duplicated at 1B3-4 inferred from genetics. new order: 1 - 1B3|20F|1B4 - 20F. origin: Rex-induced reinversion of In(1)scS1Lsc4R. references: Swanson, 1984, Ph.D. thesis, Michigan State Univ. Robbins and Swanson, 1988, Genetics 120: 1053-59. genetics: One of two mitotic exchange products induced by Rex in In(1)scS1Lsc4R [see also Dp(1;f)scS1Lsc4R]. Euchromatin and heterochromatin are restored to normal sequence except for duplication of an inverted segment of 20F containing the NO and adjacent material at 1B3-4. Presence of NO material at both ends of the chromosomes has been confirmed by examination of the bb phenotypes of the separated ends, with varied levels of bb expression observed for the ends of independently derived chromosomes. Presence of adjacent heterochromatin in the duplicated segment is inferred from the structure of In(1)scS1Lsc4R. # Dp(1;1)scS1Lsc8R cytology: Dp(1;1)1B2-3;1B3-4. origin: Associated with In(1)scS1Lsc8R. # Dp(1;1)scS1LscL8R cytology: Dp(1;1)h30-31;h32. origin: Associated with In(1)scS1Lsc8R. # Dp(1;1)scV1: see In(1LR)scV1 # Dp(1;1)SM1: Duplication (1;1) Semenova Mglinetz cytology: Dp(1;1)1A1;3E. origin: / ray induced. references: Semenova, Mglinetz, and Glotoff, 1970, Genetika (Moscow) 6(4): 165-69. Mglinetz and Semenova, 1970, Genetika (Moscow) 6(8): 86-94. # Dp(1;1)SM2 cytology: Dp(1;1)3A;11E. origin: / ray induced. references: Semenova, Mglinetz, and Glotoff, 1970, Genetika (Moscow) 6(4): 165-69. Mglinetz and Semenova, 1970, Genetika (Moscow) 6(8): 86-94. # Dp(1;1)snS93: see Tp(1;1)snS93 # *Dp(1;1)Th: Duplication (1;1) Theta origin: X-ray-induced detachment of C(1)RM with X-ray-induced deletion of most of the X euchromatin. discoverer: Muller. references: Muller and Painter, 1929, Am. Nat. 63: 197. Patterson, 1930, Genetics 15: 141-49. Muller, 1932, Proc. Intern. Congr. Genet., 6th., Vol. 1: 213-55. Stern, 1956, Wilhelm Roux's Arch. Dev. Biol. 149: 1-25. genetics: Fragment of X chromosome, including y+, sc+, and bb+ attached to right of X centromere. Causes development of interalar bristle not ordinarily present in D. melanogaster (Stern, 1956). # Dp(1;1)w: Duplication (1;1) white cytology: Dp(1;1)3A;3C. new order: 1 - 3C|3A - 20. origin: Spontaneous as a recombinant from wch/wsp. discoverer: E. B. Lewis, 55j. references: 1957, DIS 31: 84. genetics: Loci of w and rst within duplicated section. Unequal crossing over gives normal and triplicated products. Quintu- plication also produced. # Dp(1;1)w60h21: see Dp(1;1)wrG # Dp(1;1)w60h30: see Dp(1;1)wrG2 # Dp(1;1;1)w81e1 origin: Spontaneous (presumably induced by recombination between FB elements). references: Collins and Rubin, 1984, Nature 308: 323-27. genetics: Mutant for w. Male viable and fertile. molecular biology: wc insertion conserved and flanked on the left by two copies of the sequence duplicated in wi and on the right by a single copy of the sequence. # Dp(1;1;1)w82a3 origin: Spontaneous (presumably induced by recombination between FB elements). references: Collins and Rubin, 1984, Nature 308: 323-27. genetics: Mutant for w. Male viable and fertile. molecular biology: 1 kb of DNA deleted from wc inversion; oth- erwise, rearrangement same as Dp(1;1;1)w81e1. # Dp(1;1)w+1 cytology: Dp(1;1)3C1-2;4A; tandem duplication. origin: Induced by ethyl methanesulfonate. references: Green and Lefevre, 1979, Chromosoma 74: 329-35. Green, 1984, Mol. Gen. Genet. 194: 275-78. genetics: Male phenotype w+ except with z. Duplicated males fertile, but do not survive as well as wild-type males. Rev- ersion to non-duplicated chromosome in males and females. Dp(1;1)w+1, z males are z. # Dp(1;1)w+2 cytology: Dp(1;1)3C1-2;3C6-7 (probably); tandem duplication. origin: Induced by ethyl methanesulfonate. references: Green and Lefevre, 1979, Chromosoma 74: 329-35. Green, 1984, Mol. Gen. Genet. 194: 275-78. genetics: Phenotype w+. Males rarely survive and survivors are sterile; females are viable. Reversion to non-duplicated chromosome in females. Dp(1;1)w+2, z males are z (Rasmuson, Svahlin, Montell, and Olofsson, 1978, Mutat. Res. 54: 33-38). # Dp(1;1)w+61e19 cytology: Tandem duplication of proximal part of w plus tran- sposon inserted between duplications. origin: Unequal crossing over. synonym: Dp(1;1)w+R61e19; Dp(1;1)w+R; "DP" (Green, 1963). references: Green, 1963, Z. Vererbungsl. 94: 200-14. Rasmuson and Green, 1974, Mol. Gen. Genet. 133: 249-60. Rasmuson, Montell, Rasmuson, Svahlin, and Westerberg, 1980, Mol. Gen. Genet. 177: 567-70. Goldberg, Sheen, Gehring, and Green, 1983, Proc. Nat. Acad. Sci. USA 80: 5017-21. Fujikawa and Kondo, 1986, Genetics 112: 505-22. genetics: Duplication males with z are z (Rasmuson, Svahlin, Montell, and Olofsson, 1978, Mutat. Res. 54: 33-38). Dp(1;1)w+61e19, z/Dp(1;Y)y-z males show z variegation. Dupli- cation semistable, reverting to z+, the reversion occurring with a higher frequency in mei-9 than in wild-type strains (Fujikawa and Kondo, 1986). molecular biology: About 80 kb of DNA in proximal part of w duplicated (Gunaratne, Mansukhani, Lipari, Liou, Martindale, and Goldberg, 1986, Proc. Nat. Acad. Sci. USA 83: 701-05). # Dp(1;1;1)w+61e19 cytology: Tandem triplication thought to involve sub-locus 4 (or 4 and 5) of the w locus. origin: Unequal crossing over. synonym: Tr(1)w+61e19; "TR" (Green, 1963). references: Green, 1963, Z. Vererbungsl. 94: 200-14. Rasmuson and Green, 1974, Mol. Gen. Genet. 133: 249-60. Rasmuson, Montell, Rasmuson, Svahlin, and Westerberg, 1980, Mol. Gen. Genet. 177: 567-70. genetics: Triplication males with z are z. # Dp(1;1)wa: Duplication (1;1) white-apricot cytology: Dp(1;1)3A10-B1;3C3-5 (Gersh, 1962, Genetics 47: 1393-98). new order: 1 - 3C3|3B1 - 20. origin: Spontaneous from wa/wa female; recovered once as a recombinant and once as a presumed recombinant. discoverer: Green. references: 1959, Genetics 44: 1243-56. # Dp(1;1)waw+ cytology: Tandem duplication of entire w locus marked by wa in left segment and w+ in right segment. origin: Unequal crossing over in wa/w+ females. synonym: Dp(wa;w+). references: Green, 1966; Genetics 54: 881-85. 1967, Biol. Zentralbl. 86: 211-20. 1969, Mol. Gen. Genet. 103: 209-17. genetics: Male phenotype w+ except with z. # Dp(1;1)wi: Duplication (1;1) white-ivory references: Green, Todo, Ryo, and Fujikawa, 1986, Proc. Nat. Acad. Sci. USA 83: 6667-71 [except for Dp(1;1;1;1;1)wi]. genetics: Heterozygote with Dp(1;1)z shows z eye color. Eye color of wi duplication males and homozygous wi duplication females darkens with increased dosage of wi. Duplications unstable, reverting to w+. molecular biology: Quantitative increase in white locus DNA (indicated by Southern blot hybridization) as wi dosage increased. duplication cytology origin ____________________________________________________________________ Dp(1;1)wi 3A;3C; serial tandem duplication X ray Dp(1;1;1)wi serial tandem triplication unequal Xover Dp(1;1;1;1)wi ( serial tandem quadruplication unequal Xover Dp(1;1;1;1;1)wi | serial tandem quintuplication unequal Xover ( Homozygous females are sterile. | Discoverer: E.B. Lewis. Synonym: Qn(1)wi; Dp(1;1)wiQn. References: Bowman, 1969, Mutat. Res. 7: 409-15. # Dp(1;1)wm51bLwm4R (L. Robbins) cytology: Dp(1;1)20F;3C1-2. Variable portions of NO duplicated at 3C1-2; inferred from genetics. new order: 1 - 3C1|20F|3C2 - 20F. origin: Rex-induced reinversion of In(1)wm51bLwm4R. discoverer: Robbins 87g. references: Robbins and Swanson, 1988, Genetics 120: 1053-59. genetics: One of two mitotic exchange products induced by Rex in In(1)wm51bLwm4R (see also Dp(1;f)wm51bLwm4R). Euchromatin and heterochromatin are restored to normal sequence except for duplication of NO material at 3C1-2. In effect, each such chromosome is a half-tetrad containing both of the products of a single Rex-induced exchange. Examination of the bb pheno- types of the separated NO regions of a series of independently generated Dp(1;1)wm51bLwm4R chromosomes indicates that both can be mutant, both can be bb+, or either one can be mutant and the other wild type (Robbins). # Dp(1;1)wrdp cytology: Tandem duplication of the proximal part of w. references: Jack and Judd, 1978, Proc. Nat. Acad. Sci. USA 76: 1368-72. genetics: Carries wbf in distal part of w. Formed by recombi- nation between offset roo insertion elements in wric and wbf, reciprocal product is wr,def (Davis, Shen, and Judd, 1987, Proc. Nat. Acad. Sci. 84: 174-78). # Dp(1;1)wrdp+ cytology: Tandem duplication of the proximal part of w. references: Jack and Judd, 1978, Proc. Nat. Acad. Sci. USA 76: 1368-72. genetics: No wbf in distal part of w. # Dp(1;1)wrg: Duplication (1;1) white-recombinant of Green cytology: Dp(1;1)3A3-4;3C1-2 or Dp(1;1)3A4-5;3C2-3 (Gersh, 1967). new order: 1 - 3C1|3A4 - 20. origin: A regular product of asymmetric exchange between 3C1-2 of a wa or wa2 chromosome and 3A3-4 of specific homologs. Reciprocal of Df(1)wrG. discoverer: Green, 60h21. synonym: Dp(1;1)3C1; Dp(1;1)w60h21. references: 1961, Genetics 46: 1555-60. Gersh, 1962, Genetics 47: 1393-98 (fig.). 1967, Genetics 56: 309-19. Pearson, 1976, Hereditas 82: 57-62. genetics: Covers z up to and including at least part of w. # Dp(1;1)wrG2 cytology: Dp(1;1)3B2-C1;3C3-5 [Gersh, 1962, Genetics 47: 1393-98 (fig.)]. new order: 1 - 3C3|3C1 - 20. origin: Spontaneous by recombination. discoverer: Green, 60h30. synonym: Dp(1;1)w60h30. references: CP627. # Dp(1;1)wrJ2: Duplication (1;1) white-recombinant of Judd cytology: Dp(1;1)3A6-8;3C1-3 [could be same as Dp(1;1)wrG]. new order: 1 - 3C1|3A8 - 20. origin: A regular product of unequal exchange between the 3C1-3 region of a chromosome carrying wrdp with the 3A4-8 region of specific homologs. Probably reciprocal recombinant of Df(1)wrJ2. discoverer: Judd, 1961. synonym: Mutant locus designated wr,dup. references: 1961, Proc. Nat. Acad. Sci. USA 47: 545-50. # Dp(1;1)wspw17G cytology: Dp(1;1)3A1-2;3C3-4; tandem repeat of entire w locus. origin: Unequal crossing over between X marked with wsp and X marked with w17g. synonym: Dp(wsp;w17g). references: Green, 1973, Genetics Supplement 73: 187-94. Rasmuson, Green, and Karlsson, 1974, Mol. Gen. Genet. 133: 237-47. Rasmuson and Green, 1974, Mol. Gen. Genet. 133: 249-60. Rasmuson, Montell, Rasmuson, Svahlin, and Westerberg, 1980, Mol. Gen. Genet. 177: 567-70. genetics: Complementary (maroon) eye color in males resulting from interaction between markers wsp and w17g. Unstable; duplication stocks carrying z produce w+ males which, in turn, produce males with z eyes, white deficiencies, and transpositions of the w locus to other chromosomes. # Dp(1;1)ybl: Duplication (1;1) yellow-bristle cytology: Dp(1;1)1B2-3;4F8-9;5D4-5 (Nicoletti, Lindsley). new order: 1A - 1B2|5D4 - 4F9|1B3 - 20. origin: Spontaneous. discoverer: Sandler. references: Sandler, Hart, and Nicoletti, 1960, DIS 34: 103-4. genetics: Mutant for y; duplicated for cv. Regularly generates further rearrangements; has produced losses of the duplicated segment, which are accompanied by changes in phenotype from ybl to y-like and a translocation between the tips of X and 2L accompanied by a change from ybl to y+. molecular biology: Three discontinuities in DNA sequence at coordinates 40.8-41.5, 53.2-55.0, and 63.3-65.5 (Biessmann, 1985, Proc. Nat. Acad. Sci. USA 82: 7369-73). # In(1)z+: Inversion (1) zeste wild origin: X ray induced. Recovered as z reversion in genotype ordinarily exhibiting z phenotype. inversion cytology ref ( ___________________________________________ In(1)z+1 3C1;3F5-6 1 In(1)z+2 3C1-2;7A 1 In(1)z+7G3 3C3-5;14B4-C1;20A3 2 In(1)z+8 3C1-3;20D-F 1 In(1)z+11E7 3C2-5;17C4-D1;20A3 2 In(1)z+28 2B-C;3C 1 In(1)z+30 3C1-2;5D 1 In(1)z+38 3C;10C-D 1 In(1)z+48 3C1-2;4D-5D 1 In(1)z+50 3C1-2;20B-C 1 In(1)z+51 1F3-4;3B4-C1;12B-D;20 1 ( 1 = Gelbart, 1971, Ph.D. Thesis, Univ. of Wisconsin; 2 = Gans, 1953, Bull. Biol. Fr. Belg., Suppl. 38: 1-90. # Dp(1;1)z59d cytology: Dp(1;1)2F5-3A1;3A4-5 (Gersh). Tandem duplication of z. new order: 1 - 3A4|3A1 - 20. origin: X ray induced in y2 su(wa) z. discoverer: Green, 59d15. references: 1961, Genetics 46: 1555-60. Gersh, 1962, Genetics 47: 1393-98 (fig.). Jack and Judd, 1979, Proc. Nat. Acad. Sci. USA 76: 1368-72. genetics: Contains z allele but no w gene. When two doses of w+ present, z+/Dp(1;1)z females have orange mottled eyes while z+/z females have wild-type eyes (Jack and Judd, 1979). # Dp(1;Y)1Ac+: see l(1)1Ac+Y Described in section on Y derivatives. # Dp(1;Y)1E cytology: YSX.YL from which all euchromatic genes have been removed except those located in 1A through 1E. origin: X ray induced. discoverer: Masterson, 1961. synonym: T(1;Y)1E; YSsc+y.YL (Ehrlich, 1965). references: Clancy, 1964, Genetics 50: 241 (abstract). Ehrlich, 1965, Can. J. Genet. Cytol. 7: 430-32. Masterson, 1968, DIS 43: 61, 161. Lucchesi and Bischoff, 1969, Genetics 61: s37-38. genetics: Carries y and wild-type alleles of ac - su(wa). Male viable and fertile (Masterson, 1968). Variegates strongly for dor and dorl (Lucchesi and Bischoff, 1969). # Dp(1;Y)2E cytology: YSX.YL from which all euchromatic genes have been removed except those between 1A and 2B15 (Belyaeva et al., 1982). [An additional deficiency involving su(s)+ - sta+ is reported by Rayle and Hoar (1969) in a Dp(1;Y)2E obtained from Clancy.] discoverer: Masterson, 1961. synonym: T(1;Y)2E. references: Clancy, 1964, Genetics 50: 241 (abstract). Masterson, 1968, DIS 43: 61, 161. Lucchesi and Bischoff, 1969, Genetics 61: s37-38. Rayle and Hoar, 1969, DIS 44: 94. Belyaeva, Aizenzon, Kiss, Gorelova, Pak, Umbetova, Kramers and Zhimulev, 1982, DIS 58: 184-90. genetics: Carries y and wild-type alleles of ac and dor or l(1)2Bh; possibly deficient for su(s) - sta. Male viable and fertile. Slight variegation for dor (Lucchesi and Bischoff, 1969). # Dp(1;Y)59k9-1 cytology: Dp(1;Y)2A2-B1. references: Rayle and Hoar, 1969, DIS 44: 94. genetics: Carries sta+. # Dp(1;Y)60e17.4-3 cytology: Dp(1;Y)1E2-4. references: Rayle and Hoar, 1969, DIS 44: 94. genetics: Carries y+ - su(wa)+. # Dp(1;Y)68h20 cytology: Dp(1;Y)2B3-5. references: Rayle and Hoar, 1969, DIS 44: 94. genetics: Carries y+ - sta+. # Dp(1;Y)BSv+y+: see BSv+Yy+ Described in section on Y derivatives. # Dp(1;Y)BSw+: see BSw+Y Described in section on Y derivatives. # Dp(1;Y)BSw+y+: see BSw+y+Y Described in section on Y derivatives. # Dp(1;Y)BSy61d: see BSYy61d Described in section on Y derivatives. # Dp(1;Y)BSy67j: see BSYy67j Described in section on Y derivatives. # Dp(1;Y)BSybl2: see BSYybl2 Described in section on Y derivatives. # Dp(1;Y)BS-Y: see v+Yy+3 Described in section on Y derivatives. # Dp(1;Y)BSV: see BSVY Described in section on Y derivatives. # Dp(1;Y)bw+: see bw+Y Described in section on Y derivatives. # Dp(1;Y)bw+y+: see bw+Yy+ Described in section on Y derivatives. # Dp(1;Y)cin+: see cin+Y Described in section on Y derivatives. # Dp(1;Y)ct+y+: see ct+y+Y Described in section on Y derivatives. # Dp(1;Y)dor: Duplication (1;Y) deep orange origin: / ray induced in attached-XY, dor males. discoverer: Gorelova. synonym: dorY. references: Belyaeva, Aizenzon, Kiss, Gorelova, Pak, Umbetova, Kramers, and Zhimulev, 1982, DIS 58: 184-90. duplication cytology genetics __________________________________________ Dp(1;Y)dor1T 1A;2C1-2 Dp(1;Y)dor3T 1A;2E1-2 Dp(1;Y)dor6T y+-arm+ Dp(1;Y)dor9T y+-hfw+ Dp(1;Y)dor10T 1A;2C1-2 Dp(1;Y)dor13T 1A;2B9-10 y+-hfw+ Dp(1;Y)dor17T 1A;2C1-2 y+-arm+ Dp(1;Y)dor18T 1A;2D1-2 Dp(1;Y)dor21T ( 1A;2B1-2 + 3 thick bands ( The three thick condensed bands pair with the 2B3-4 - 2B9-10 bands of the wild type X. No 2B5 puff in this duplication. # Dp(1;Y)dor+: see dor+Yy+ Described in section on Y derivatives. # Dp(1;Y)l(1)J1+: see l(1)1Ac+Y Described in section on Y derivatives. # Dp(1;Y)mal2y+: see y+Ymal+2 Described in section on Y derivatives. # Dp(1;Y)mal+ cytology: Dp(1;Y)19A1;20F (Eberl). new order: YL|19A1 - 20F|YS. origin: X-ray-induced deletion of majority of euchromatin [l(1)J1+ through car+] from YSX.YL, In(1)EN. discoverer: E.H. Grell. synonym: Ymal+. references: Brosseau, Nicoletti, Grell, and Lindsley, 1961, Genetics 46: 339-46. Rahman and Lindsley, 1981, DIS 46: 339-46. Eberl and Hilliker, 1988, Genetics 118: 109-20. genetics: Carries sw+-bb+. # Dp(1;Y)pn-w+: see pn-w+Y Described in section on Y derivatives. # Dp(1;Y)scS1: see YL.uscS1 Described in section on Y derivatives. # Dp(1;Y)scV1: see scV1.YS+ Described in section on Y derivatives. # Dp(1;Y)shi+: see shi+Y Described in section on Y derivatives. # Dp(1;Y)su(f)-: see su(f)-Y Described in section on Y derivatives. # Dp(1;Y)Sz: Duplication (1;Y) Szeged origin: X ray induced in Oregon-R (premeiotic?). discoverer: Koczka and Kiss. synonym: YSz. references: Belyaeva, Aizenzon, Kiss, Gorelova, Pak, Umbetova, Kramers, and Zhimulev, 1982, DIS 58: 184-90. duplication X cytology genetics ______________________________________________________ Dp(1;Y)Sz71/4 1A;2C1-2 y+-arm+ Dp(1;Y)Sz78/4 1A;2B17-18 y+-arm+ Dp(1;Y)Sz170 1A;2B17-18 y+-arm+ Dp(1;Y)Sz280 1A;2C1-2 + y+-sta+, BRC--hfw-, Df(1)2B3-4;2B7-8 l(1)Bg+-arm+ Dp(1;Y)Sz303 1A;2B17-18 y+-arm+ # Dp(1;Y)v+y+: see v+Yy+ Described in section on Y derivatives. # Dp(1;Y)w+: see w+Y Described in section on Y derivatives. # Dp(1;Y)w+303 cytology: Dp(1;Y)2D1-2;3D3-4. origin: X ray induced. discoverer: Lefevre. references: Perrimon, Engstrom, and Mahowald, 1984, Genetics 108: 559-72. 1985, Genetics 111: 23-41. genetics: Carries w+ and wild-type alleles of loci distal and proximal to w+. # Dp(1;Y)w+y+: see w+y+Y Described in section on Y derivatives. # Dp(1;Y)y2: Duplication (1;Y) yellow origin: / ray induced in attached -XY, y2 su(wa) wa males. discoverer: Gorelova. synonym: y2Y. duplication cytology ref ( genetics ________________________________________________ Dp(1;Y)y221T 1A;2B6(2B7-8?) 1,2 y+-hfw+ Dp(1;Y)y222T 1A;2B9-10 1 y+-fmf+ Dp(1;Y)y240T 1A;2B9-11 1,2 y+-fmf+ Dp(1;Y)y243T 1A;2E1-2 1 Dp(1;Y)y253T 1A;2B6(2B7-8?) 1,2 y+-hfw+ ( 1 = Belyaeva, Aizenzon, Kiss, Gorelova, Pak, Umbetova, Kra- mers, and Zhimulev, 1982, DIS 58: 184-90; 2 = Belyaeva, Aizenzon, Semeshin, Kiss, Koczka, Baritcheva, Gorelova, and Zhimulev, 1980, Chromosoma 81: 281-306. # Dp(1;Y)y261l: see y2Y61l Described in section on Y derivatives. # Dp(1;Y)y267g: see y2Y67g Described in section on Y derivatives. # Dp(1;Y)y2sc cytology: Dp(1;Y)1E-F;2B-C;3A. new order: 1A1 - 1E-F|2B-C - 3A. references: Eberl and Hilliker, 1988, Genetics 118: 109-20. # Dp(1;Y)y59b2: see y59bY2 Described in section on Y derivatives. # Dp(1;Y)y+mal1: see y+Ymal+1 Described in section on Y derivatives. # Dp(1;Y)y+mal2: see y+Ymal+2 Described in section on Y derivatives. Dp(1;Y)y+mal+102 - Dp(1;Y)y+mal+126: see y+Ymal+102 - y+Ymal+126 Described in section on Y derivatives. # Dp(1;Y)y+sc: see y+scY Described in section on Y derivatives. # Dp(1;Y)y-z: Duplication (1;Y) yellow-zeste references: Gunaratne, Mansukhani, Lipari, Liou, Martindale, and Goldberg, 1986, Proc. Nat. Acad. Sci. USA 83: 701-05. genetics: Contains wild-type alleles of gt and tko but not l(1)3Ac. Dp(1;1)w+61e19/Dp(1;Y)y-z males show z variegation. molecular biology: X breakpoint localized to an approximately 4-kb restriction fragment 7-8 kb to the right of the origin of a 160-kb walk in the vicinity of z. # Dp(1;Y;2)BS5 cytology: Piece of Y carrying BS attached to 2L .01 unit to left of locus of al. origin: X ray induced in BSYy+. references: Novitski, Ehrlich, and Becker, 1971, DIS 47: 91- 92. genetics: Carries BS and fertility factor kl-5; claimed to carry ks-1 and possibly kl-4 (= kl-3?) as well. # Dp(1;Y;2)bw+: see X.bw+YL Described in section on XY combinations. # Dp(1;Y;3)H: see scV1.YSmwh+ Described in section on Y derivatives. # Dp(1;Y:3)M1-3 cytology: Duplication of 61 to 62A-B, including mwh+, on tip of YSX chromosome distal to the KS factors. origin: X ray induced in T(Y;3)P6, mwh+ ve+/ y+YSX., y+KS y cv v f; mwh ve males. synonym: mwh+YSX. references: Garcia-Bellido and Ripoll, 1973, DIS 50: 92. genetics: Male viable and fertile over YL chromosome. Homozy- gous females lethal. No variegation for the mwh+ of the duplication in nullo-Y males. # Dp(1;Y;4)BS origin: X ray induced in BSYy+/C(1)RM, y2 su(waa) wa bb females. references: Parker, 1965, Mutat. Res. 2: 523-29. genetics: Y carries BS and shows Y-4 linkage. # Dp(1;Y;4)y+ origin: X ray induced in BSYy+/C(1)RM, y2 su(wa) wa bb females. references: Parker, 1965, Mutat. Res. 2: 523-29. genetics: Y carries y+ and shows Y-4 linkage. # Dp(1;2)51b: see Tp(1;2)51b # Dp(1;2)51bV76e cytology: Dp(1;2)3C1-2;3D6-E1;40-41;52E. new order: 21 - 41|3C2 - 3D6|52E - 41|52E - 60 or 21 - 40|52E - 40|3C2 - 3D6|52E - 60. origin: Spontaneous. references: Poulson and Lefevre, 1982, Genetics 100: s54-55. genetics: When duplication covers l(1)NB, N55e, N60g11, N264- 40, N264-103, or NNic, some males show variegated or roughened eyes, abnormal legs, missing or multiple bristles, and missing or fused ocelli. # Dp(1;2)63i: see Dp(1;2)v+63i # Dp(1;2)65b: see Tp(1;2)v65b # Dp(1;2)76f cytology: Deletion of N+ to dm+ region in Dp(1;2)51b. Retains functions of w+ - rst+. origin: Induced by ethyl methanesulfonate. references: Welshons and Welshons, 1985, Genetics 110: 465-77. 1986, Genetics 113: 337-54. genetics: Covers w, rst, and vt but not N. # Dp(1;2)(w-ec)64d: see Tp(1;2)w-ec # Dp(1;2;3)pn3: see Tp(1;2;3)pn3 #*Dp(1;2)A12: see *Tp(1;2)A12 #*Dp(1;2)A124: see *Tp(1;2)A124 # Dp(1;2)BS cytology: Second chromosome carrying fragment of BSYy+ with the marker BS attached to 2L tip. origin: X ray induced in female with BSYy+. references: Novitski, Ehrlich, and Becker, 1971, DIS 47: 91- 92. Novitski, 1976, The Genetics and Biology of Drosophila (Ash- burner and Novitski, eds.). Academic Press, London, New York, San Francisco, Vol. 1b., pp. 562-68. # Dp(1;2)BS3 cytology: BS attached to 2L 0.1 unit to left of locus of al. origin: X ray induced in BSYy+. references: Novitski, Ehrlich, and Becker, 1971, DIS 47: 91- 92. genetics: Homozygous viable and fertile. Carries BS but no male fertility factors; probably only X-derived material translocated to 2L. #*Dp(1;2)ct7c1: see *Tp(1;2)ct7c1 # Dp(1;2)E1 cytology: Dp(1;2)1A-F;60A. references: Jacobson, Yim, Grell, and Wobbe, 1982, Cell 30: 817-23. genetics: Carries normal alleles of y+ and su(s)+. # Dp(1;2)f+ cytology: f+ inserted near centromere of 2. origin: X ray induced. references: Shukla and Auerbach, 1980, Genet. Res. 36: 41-56. genetics: Variegates for f. Transposes to other chromosome sites on 2 or 4. # Dp(1;2)FN107: see Tp(1;2)sn+72d # Dp(1;2)HwK cytology: Dp(1;2)1B2-3;1E-F;21C2-3. new order: 21A - 21C2|1B3 - 1E|21C3 - 60. discoverer: Kreber. references: Craymer. genetics: Mutant for Hw. # Dp(1;2)K1: Duplication (1;2) of Krivshenko cytology: Dp(1;2)1B1-3;20;29A-B; deficient for 1B3-20. Larval ganglion cells in metaphase show a rod-shaped element, believed to be distal 2L capped with the X centromere, and a J-shaped element, believed to be the X tip attached to the rest of 2 with the 2 centromere. new order: 1A1 - 1B1|29B - 60; 20|29A - 21. origin: X ray induced in Canton-S wild type. synonym: T(1;2)K1. references: Krivshenko, 1956, DIS 30: 74-76. genetics: Bipartite duplication, the 2PXD segment carrying y+ and the XP2D segment carrying bb+. Flies hyperploid for the XP2D segment survive; flies with a normal X but homozygous for Dp(1;2)K1 die. # Dp(1;2)Nst: see Tp(1;2)w-ec # Dp(1;2)pn1: see Tp(1;2)pn1 # Dp(1;2)pn-ec: see Tp(1;2)w62g26 # Dp(1;2)r+75c: see Tp(1;2)r+75c # Dp(1;2)rb+71g: see Tp(1;2)rb+71g # Dp(1;2)sc19: see Tp(1;2)sc19 # Dp(1;2)sc260-27: see Tp(1;2)sc260-27 # Dp(1;2)sn+72d: see Tp(1;2)sn+72d # Dp(1;2)v65b: see Tp(1;2)v65b # Dp(1;2)v+63i cytology: Dp(1;2)9E1;10A11;56A. origin: X ray induced. discoverer: Lefevre. synonym: Dp(1;2)63i; Tp(1;2)v+63i. references: 1969, Genetics 63: 589-600. Hall and Kankel, 1976, Genetics 83: 517-35. Craymer and Roy, 1980, DIS 55: 200-04. Lefevre, 1981, Genetics 99: 461-80. Zhimulev, Belyaeva, Pokholkova, Kotchneva, Fomina, Bgatov, Khudyakov, Patzevich, Semeshin, Baritcheva, Aizenzon, Kra- mers, and Eeken, 1981, DIS 56: 192-96. genetics: Covers ras - l(1)10Ag, including v. RpII215 not included. # Dp(1;2)v+75d: see Tp(1;2)v+75d # Dp(1;2)w51b: see Tp(1;2)51b # Dp(1;2)w+51b7: see Tp(1;2)51b # Dp(1;2)w+62g26: see *Tp(1;2)w+62g26 # Dp(1;2)w+70h cytology: Dp(1;2)3A7-8;3C2-3;31A3. discoverer: Green. references: Judd, Shen, and Kaufman, 1972, Genetics 71: 139- 56. Jack and Judd, 1979, Proc. Nat. Acad. Sci. USA 76: 1368-72. genetics: Covers mit(1)15 - w. molecular biology: Right breakpoint some 28 kb to the right of the wa copia insertion point (Pirrotta, Hadfield, and Pre- torius, 1983, EMBO J. 2: 927-34). #*Dp(1;2)wm52b: see *Tp(1;2)wm52b #*Dp(1;2)wm53a: see *Tp(1;2)wm53a #*Dp(1;2)wm258-44: see *Tp(1;2;3)wm258-44 # Dp(1;2)w-ec: see Tp(1;2)w-ec # Dp(1;2;3)pn3: see Tp(1;2;3)pn3 # Dp(1;3)51 cytology: X broken distally and tip attached to centric part of 3; also broken proximally and centromere of X attached to acentric part of 3. Interstitial part of X without centromere lost. Third-chromosome breakpoint not determined. origin: X ray induced. discoverer: Weltman. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. genetics: Bipartite duplication, the 3PXD segment carrying sc+-su(wa)+ and the XP3D segment carrying bb+. # Dp(1;3)126 origin: X ray induced. discoverer: Dobzhansky, 1930. references: 1935, Z. Indukt. Abstamm. Vererbungsl. 68: 143. genetics: Duplicated for r, M(1)15D, f, and B but not sl or os; variegates for f and M(1)15D (Schultz). Duplicated section inserted into chromosome 3 between st and cu. Also an inver- sion in 3L. Viability, fertility, and phenotype of Dp(1;3)126/+ male and female normal. # Dp(1;3)142 cytology: X broken distally and tip attached to centric part of 3; also broken proximally and centromere of X attached to acentric part of 3. Interstitial part of X without centromere lost. Third-chromosome breakpoint not determined. origin: X ray induced. discoverer: Weltman. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. genetics: Bipartite duplication, the 3PXD segment carrying sc+-su(wa)+ and the XP3D segment carrying su(f)+-bb+. # Dp(1;3)BS cytology: Duplication with BS and some heterochromatin on 3L tip; BS located to left of ru. origin: X ray induced in BSYy+. discoverer: Leigh. references: Novitski, Grace, and Strommen, 1981, Genetics 98: 257-73. Puro and Novitski, 1982, DIS 58: 126. genetics: Carries BS. Female homozygotes viable and fertile, male homozygotes viable but sterile. Used in construction of C(3)EN (Novitski et al., 1981). # Dp(1;3)BS3i: see Tp(1;3)BS3i #*Dp(1;3)ct11a: see *Tp(1;3)ct11a #*Dp(1;3)ct12c1: see *Tp(1;3)ct12c1 #*Dp(1;3)ct268-37: see *Tp(1;3)ct268-37 # Dp(1;3)ctJ8: see Tp(1;3)ctJ8 # Dp(1;3)Dspot: see Tp(1;3)N264-58 # Dp(1;3)E1 cytology: Dp(1;2)1A-F;61A. references: Jacobson, Yim, Grell, and Wobbe, 1982, Cell 30: 817-23. genetics: Carries normal alleles of genes from the tip of X through brc (Voelker). # Dp(1;3)f+71b: see Tp(1;3)f+71b # Dp(1;3)Gspot: see Dp(1;3)w67k27 # Dp(1;3)in61j2: Duplication (1;3) inturned cytology: Dp(1;3)20;77B-C. Nucleolus moved to in region. origin: X ray induced. references: Arajarvi and Hannah-Alava, 1969, DIS 44: 73-74. Hannah-Alava, 1971, Mol. Gen. Genet. 113: 191-203. genetics: Homozygous lethal, and lethal with some in alleles induced by X ray. Expression of in variable with other in alleles, becoming more nearly wild type with addition of a Y (Hannah-Alava, 1971). # Dp(1;3)JC153: see Tp(1;3)JC153 # Dp(1;3)K2: Duplication (1;3) of Krivshenko cytology: Dp(1;3)20A-B;20D-F;80-81 superimposed on In(1)1B2- 3;20B-D1. Inferred from genetic data since salivary chromo- somes do not reveal an aberration. In ganglial metaphase, chromosome 3 is a rod-shaped and a J-shaped element. new order: 1A - 1B2|20B|80 - 100; 20F|80 - 61. Tentative. origin: X ray induced in In(1)sc8. synonym: T(1;3)K2. references: Krivshenko, 1956, DIS 30: 76. genetics: Bipartite duplication. Irradiated In(1)sc8 broken in distal region between y+ and bb+ and also near the centromere. Chromosome 3 broken near the centromere, whether to left or right of the centromere is not known. Tip of X chromosome with y+ and ac+ is attached to the chromosome 3 centromere, and one arm of this chromosome is attached to the X cen- tromere. Bulk of the X chromosome is thus acentric and lost. Homozygote viable and moderately fertile. # Dp(1;3)Lspot: see Dp(1;3)wm49a # Dp(1;3)m+84f cytology: Dp(1;3)10B;10F-11A;98. new order: 61 - 98|10B - 10F|98 - 100. origin: Spontaneous. discoverer: Voelker. genetics: Covers Df(1)m259-4 and therefore includes wild-type alleles of tyl and RpII215 (Voelker). Exists only as the duplication (deficiency segregant lost). # Dp(1;3)MNB-8 cytology: Breaks near the two 16F regions of Dp(1;1)MNB-8 with resulting acentric fragment inserted between 67E1 and 67E7 of 3; deficient X lost. new order: 61A - 67E|16F - 16A1|16A7-16A1|17E1 - 17A1|67E - 100. origin: Spontaneous derivative of Dp(1;1)MNB-8. references: Holmquist, 1972, Chromosoma 36: 413-52. genetics: Carries B os+. # Dp(1;3)N50k: see Tp(1;3)N50k # Dp(1;3)N264-58: see Tp(1;3)N264-58 #*Dp(1;3)N264-100: see *Tp(1;3)N264-100 # Dp(1;3)O4: see Tp(1;3)O4 # Dp(1;3)pn25: see Tp(1;3)pn25 # Dp(1;3)pn26: see Tp(1;3)pn26 # Dp(1;3)rasv: see Tp(1;3)rasv #*Dp(1;3)sc260-20: see *T(1;3)sc260-20 # Dp(1;3)scJ4: see T(1;3)scJ4 # Dp(1;3)sn13a1: see Tp(1;3)sn13a1 # Dp(1;3)sta: see Tp(1;3)sta # Dp(1;3)v+74c: see Tp(1;3)v+74c #*Dp(1;3)w+54a4: see *Tp(1;3)w+54a4 #*Dp(1;3)w+54c10: see *Tp(1;3)w+54c10 # Dp(1;3)w+67k: see Tp(1;3)w+67k # Dp(1;3)whalo: see Tp(1;3)whalo # Dp(1;3)wm49a: see Tp(1;3)wm49a # Dp(1;3)wm264-58: see Dp(1;3)N264-58 # Dp(1;3)wvco: see Tp(1;3)wvco # Dp(1;3)wzh: see Tp(1;3)wzh # Dp(1;3)z7: see T(1;3)z7 # Dp(1;3;4)7 cytology: Dp(3;4)62E;78D;79F;102F + Dp(1;3)1B;61A from T(1;3)scJ4 new order: 1A - 1B|61A - 62F|(78D - 79F)|102F - 101. references: Duncan and Lewis, 1982, Symp. Soc. Dev. Biol., 40th, pp. 533-54. phenotype: Covers homozygous lethal Pc mutants. # Dp(1;4)20G1L cytology: Duplication for bands 1A1 to 3C2 translocated to 4. 3C1 only may be included (Gans, 1953) or 3C1 and 3C2 (Gersh, 1967). references: Gans, 1953, Bull. Biol. Fr. Belg. (Suppl.) 38: 1- 90. Gersh, 1967, Genetics 56: 309-19. # Dp(1;4)174 cytology: Duplication linked with 4 but cytology not worked out. origin: X ray induced. discoverer: Weltman. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. genetics: Probably bipartite duplication like Dp(1;3)51. Car- ries sc+ but not bb+. # Dp(1;4)193 cytology: X tip attached to 4 at left end. origin: Derived from Dp(1;4)1021. synonym: y+.spapol. references: Williamson, Parker, and Manchester, 1970, Mutat. Res. 9: 287-97, 299-306. Parker, 1970, Mutat. Res. 9: 307-32. genetics: Carries y+ and ac+ as well as ci+, ey+, and spapol. (Williamson et al., 1970). Shows 66% segregation from the compound X in C(1)RM females without a Y. # Dp(1;4)1021 cytology: Heterochromatic left arm of 4 capped by euchromatic tip of X. references: Parker, 1969, Mutat. Res. 7: 393-407. Parker and Busby, 1973, Mutat. Res. 18: 33-46. O'Tousa, 1982, Genetics 102: 503-04. genetics: Carries y+ and ac+. #*Dp(1;4)A1: see *Tp(1;4)A1 #*Dp(1;4)A12: see *Tp(1;4)A12 # Dp(1;4)exd+: Duplication extradenticle wild type discoverer: D. Falk. references: Wieschaus, Nusslein-Volhard, and Jurgens, 1984, Wilhelm Roux's Arch. Dev. Biol. 193: 296-307. genetics: Covers exd. duplication cytology synonym __________________________________________________________ Dp(1;4)exd81h24b 13F;14B5-18;14E1-4;16A2 Dp(1;4)81h24b Dp(1;4)exd82b26c 13F;14D1-2;15A3-5;16A2 Dp(1;4)82b26c other information: Also described as Df(1)81h24b and Df(1)82b26c. # Dp(1;4)mg cytology: Dp(1;4)2B7-C1;3C1-2;102F. discoverer: Green. references: Robbins, 1977, Genetics 87: 655-84. 1980, Genetics 96: 187-200. 1984, Genetics 108: 361-75. genetics: Covers lethals between z and w; includes pn+ but is w or w-. Dp(1;4)mg hyperploids have reduced viability and homozygotes having an otherwise normal genotype are lethal. Homozygotes survive if the fly is heterozygous, homozygous or hemizygous for deficiencies in the z-w region. Viability of Dp(1;4)mg hyperploids improves at lower temperatures and in XO males, suggesting position-effect-variegation of the tran- sposed genes (Robbins). #*Dp(1;4)N264-85: see *Tp(1;2;4)N264-85 #*Dp(1;4)N264-86: see *Tp(1;4)N264-86 # Dp(1;4)r+: Duplication (1;4) rudimentary - wild type cytology: Dp(1;4)14A1-2;16A7-B1;102F2-3. origin: X-ray-induced derivative of T(1;4)BS = T(1;4)15F9- 16A1;16A7-B1;102F2-3 of most of the X euchromatin from the 4PXD element deleted; likely to carry genes from X terminus. new order: 1A|14A2 - 16A7|102F2 - 101. discoverer: Green. synonym: Dp(1;4)r+f+. references: 1963, Genetica 34: 242-53. Jarry, 1979, Mol. Gen. Genet. 172: 199-202. Craymer and Roy, 1980, DIS 55: 200-04. Naguib and Jarry, 1981, Genet. Res. 37: 199-207. Ganetzky and Wu, 1982, Genetics 100: 597-614. Falk, Roselli, Curtis, Holliday, and Klufas, 1984, Mutat. Res. 126: 25-34. Ganetzky, 1984, Genetics 108: 897-911. Kulkarni, Steinlauf, and Hall, 1988, Genetics 118: 267-85. genetics: Carries normal alleles of exd, para, r and f appended to the right end of chromosome 4. Usually lethal in two doses. # Dp(1;4)wm5: see T(1;4)wm5 #*Dp(1;4)wm51c: see *Tp(1;4)wm51c # Dp(1;4)wm65g cytology: Dp(1;4)3B1-2;3C4-5;101. origin: X ray induced. references: Lefevre, 1968, DIS 43: 62-63, 165. Judd, Shen, and Kaufman, 1972, Genetics 71: 139-56. Robbins, 1980, Genetics 96: 187-200. 1984, Genetics 108: 361-75. genetics: Contains wild-type alleles of l(1)3Bb-rst. Covers Df(1)wm4Lrst3R and Df(1)w64d8 lethality. Df(1)w258-45;Dp(1;4)wm65g male zygotes are lethal in the absence of a Y. Duplication shows variegation, XO males car- rying w and Dp(1;4)m65g having almost white eyes (Robbins, 1980). # Dp(1;4)wvD1: Duplication (1;4) white-variegated of Demerec cytology: Dp(1;4)3C1-4;101A-D. origin: X ray induced in y. discoverer: Demerec, 33j19. references: CP627. genetics: Variegated for w but not ci. X broken between w and rst; 4 probably broken in left arm. #*Dp(1;f)1: Duplication (1;free) origin: X-ray-induced deletion of most of X euchromatin. discoverer: Muller. synonym: Del(1)1. references: Painter and Muller, 1929, J. Heredity 20: 287-98. Muller and Painter, 1932, Z. Indukt. Abstamm. Vererbungsl. 62: 316-65. genetics: Contains wild-type alleles of y, sc, br, pn, and bb. #*Dp(1;f)2 origin: X-ray-induced deletion of most of X euchromatin. discoverer: Muller. synonym: Del(1)2. references: Painter and Muller, 1929, J. Heredity 20: 287-98. Muller and Painter, 1932, Z. Indukt. Abstamm. Vererbungsl. 62: 316-65. genetics: Contains wild-type alleles of y, sc, br, and bb. # Dp(1;f)3 cytology: Dp(1;f)1D4-E1;20D. 4 times the size of chromosome 4 at metaphase. new order: 1A - 1D3|20A - 20F. origin: X-ray-induced deletion of most of X euchromatin. discoverer: Weltman, 1954. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. Grell, 1964, Genetics 50: 151-66. Rayle and Hoar, 1969, DIS 44: 94. Schalet and Lefevre, 1973, Chromosoma 44: 183-200. Sandler and O'Tousa, 1979, Genetics 91: 537-51. genetics: Carries wild-type alleles of y-tw and wap-bb . Includes ABO-X (Parry and Sandler, 1974, Genetics 77: 535- 39). Disjoins regularly from XY, shows 3% nondisjunction from C(1)RM, and causes 18% nondisjunction of In(1)dl-49 from + in In(1)dl-49/+/Dp(1;f)3 female. # Dp(1;f)3bb- cytology: Dp(1;f)1D3-E1;20A-F. 2 times size of chromosome 4 at metaphases; lacks NO. origin: X ray induced in Dp(1;f)3. discoverer: Parry. references: Sandler and O'Tousa, 1979, Genetics 91: 537-51. genetics: Deficient for bb. #*Dp(1;f)10 origin: X-ray-induced deletion of most of X euchromatin. discoverer: Weltman, 1954. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. genetics: Carries wild-type alleles of y, ac, and sc but not su(f) or bb. Sixty-one percent nondisjunction from XY, 45% from C(1)RM, and regular disjunction of + from In(1)dl-49 in In(1)dl-49/+/Dp(1;f)10 female. #*Dp(1;f)12 cytology: 3.4-4 times the size of chromosome 4 at metaphase; lacks only the distalmost part of heterochromatic segment hD (Cooper). origin: X-ray-induced deletion of most of X euchromatin. discoverer: Weltman, 1954. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. genetics: Contains wild-type alleles of y, ac, sc, su(wa), su(f), and bb but not pn or car. Disjoins regularly from XY, shows 4% nondisjunction from C(1)RM, and causes 19% nondis- junction of + from In(1)dl-49 in In(1)dl-49/+/Dp(1;f)12 female. #*Dp(1;f)14 origin: X-ray-induced deletion of most of X euchromatin. discoverer: Muller. synonym: Del(1)14. references: Painter and Muller, 1929, J. Heredity 20: 287-98. Muller and Painter, 1932, Z. Indukt. Abstamm. Vererbungsl. 62: 316-65. genetics: Contains wild-type alleles of y, sc, and bb but not br. # Dp(1;f)18 cytology: Dp(1;f)1E4-F1;19-20. origin: X-ray-induced deletion of most of X euchromatin. discoverer: Weltman, 1954. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. Rayle and Hoar, 1969, DIS 44: 94. Belyaeva, Aizenzon, Kiss, Gorelova, Pak, Umbetova, Kramers, and Zhimulev, 1982, DIS 58: 184-90. genetics: Contains wild-type alleles of y-l(1)1Ec and bb but not l(1)Ed (Voelker) or car. # Dp(1;f)24 cytology: Dp(1;f)1B5;19-20 (Lefevre). origin: X-ray-induced deletion of most of X euchromatin. discoverer: Muller. synonym: Del(1)24. references: 1932, Proc. Intern. Congr. Genet., 6th., Vol. 1: 213-55. Garcia-Bellido, 1979, Genetics 91: 491-520. White, Decelles, and Endow, 1983, Genetics 104: 433-48. genetics: Mutant for sc and carries wild-type alleles of l(1)1Ac, y, ac, and l(1)Bb but not l(1)Ec. # Dp(1;f)52 cytology: Dp(1;f)1B10-C4;19-20 (Gersh). 3.7-4 times the size of chromosome 4 at metaphase; lacks only the distalmost part of heterochromatic segment hD (Cooper). origin: X-ray-induced deletion of X euchromatin. discoverer: Weltman, 1954. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. Schalet and Finnerty, 1968, DIS 43: 128-29. genetics: Contains wild-type alleles of y, ac, sc, su(f), and bb but not su(wa), pn, or car. Segregates normally from XY, shows 3% nondisjunction from C(1)RM, and causes 13% nondis- junction of + from In(1)dl-49 in In(1)dl-49/+/Dp(1;f)52. # Dp(1;f)60g origin: A spontaneous exchange between the distally located heterochromatin of In(1)sc8, y31d, and the proximal hetero- chromatin of a normal X. Occurred in a triploid female. discoverer: Mohler, 60g. references: 1960, DIS 34: 52. genetics: Carries y31d, ac+, and probably two copies of su(f)+ but not car+. other information: The reciprocal product, a reversed acrocen- tric compound X [C(1)RA60g] was recovered from the same fly. # Dp(1;f)65XC2 origin: X ray induced in R(1)2 males. discoverer: Thompson, 1965. references: Gethman, 1967, DIS 42: 39, 70. genetics: Carries wild-type alleles of y, ac, sc, and su(f) but not pn or car. Covers deficiency of C(1)RA60g. Somatically unstable; mosaics frequent. Lethal in 30% of the zygotes. #*Dp(1;f)100 cytology: Two-thirds the length of normal X at metaphase. origin: Spontaneous deletion of most of X euchromatin. discoverer: L. V. Morgan, 22l. synonym: sc-Dp. references: 1938, Genetics 23: 423-62. genetics: Contains wild-type alleles of y-pn and fu-bb. Pheno- type of duplication-bearing female nearly wild type, but occipital bristles and hairs are present, eyes are a trifle smaller and rougher, and wings have straighter outer margins and sometimes scalloped inner margins. In male, duplication more than 99% lethal. # Dp(1;f)101 cytology: Dp(1;f)2A2;20E-F (Lefevre); Df(1;f)2B1-2;19F5-20A (Belyaeva et al., 1982); one-fourth the length of normal X at metaphase. new order: 1A - 2A2|20F. origin: X-ray-induced deletion of most of X euchromatin. discoverer: Dobzhansky, 1930. references: 1932, Tr. Lab. Genet. (Leningrad) 9: 193-216. 1935, Z. Indukt. Abstamm. Vererbungsl. 68: 134-62. Rayle and Hoar, 1969, DIS 44: 95. Belyaeva, Aizenzon, Kiss, Gorelova, Pak, Umbetova, Kramers, and Zhimulev, 1982, DIS 58; 184-90. genetics: Contains wild-type alleles of y-l(1)1Fc and bb but not l(1)1Fd (Voelker) or su(f). With duplication, both sexes viable and wild type except for presence of occipital bris- tles. molecular biology: DNA breakpoint (2B1-2) between coordinates 31.3 and 35.3 (Chao and Guild, 1986, EMBO J. 5: 143-50). #*Dp(1;f)102 cytology: One-fifth the length of normal X at metaphase. origin: X-ray-induced deletion of most of X euchromatin. discoverer: Dobzhansky, 1930. references: 1932, Biol. Zentralbl. 52: 493-509. 1935, Z. Indukt. Abstamm. Vererbungsl. 68: 134-62. genetics: Contains y+ to rb+ inclusive and not bb+. Usually male lethal, but female survives and has occipital bristles, narrow parallel-sided wings, branched posterior crossveins, and heavier bristles on thorax. #*Dp(1;f)106 cytology: Metaphase length about 4 times that of chromosome 4. origin: X-ray-induced deletion of most of X euchromatin. discoverer: Dobzhansky, 1930. references: 1932, Biol. Zentralbl. 52: 493-509. genetics: Contains wild-type alleles of y, sc, and svr but not bb. # Dp(1;f)107 cytology: Metaphase length about one-fifth that of a normal X. origin: X-ray-induced deletion of most of X euchromatin. discoverer: Dobzhansky, 1930. references: 1932, Biol. Zentralbl. 52: 493-509. genetics: Contains wild-type alleles of y, sc, svr, su(s) (Voelker) and bb. # Dp(1;f)112 cytology: Dp(1;f)1E4-F1;19-20 (Gersh); slightly longer than chromosome 4 at metaphase. origin: X-ray-induced deletion of most of X euchromatin. discoverer: Dobzhansky, 1930. references: 1932, Biol. Zentralbl. 52: 493-509. Belyaeva, Aizenzon, Kiss, Gorelova, Pak, Umbetova, Kramers, and Zhimulev, 1982, DIS 58: 184-90. genetics: Contains wild-type alleles of y-l(1)1Ec but not l(1)1Ed or bb. Both sexes viable and have occipital bristles. # Dp(1;f)118 cytology: About one-fourth the length of normal X at metaphase. origin: X-ray-induced deletion of most of X euchromatin. discoverer: Dobzhansky, 1930. references: 1932, Biol. Zentralbl. 52: 493-509. genetics: Contains wild-type alleles of y, sc, svr, su(s) (Voelker) and bb but not kz. # Dp(1;f)118YM cytology: About size of chromosomes 4 at metaphase. origin: Presumably same as Dp(1;f)118. references: Yamamoto and Miklos, 1977, Chromosoma 60: 283-96. genetics: Contains wild-type alleles of y, sc, and svr but not bb (Yamamoto and Miklos, 1977). # Dp(1;f)122 cytology: Dp(1;f)1E4-F1;19-20 (Gersh). origin: X-ray-induced deletion of most of X euchromatin. discoverer: Weltman, 1954. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. La Volpe, La Mantis, Gargiulo, and Malva, 1984, Mol. Gen. Genet. 194: 485-88. genetics: Carries wild-type alleles of y, ac, sc, and bb but not su(wa), pn, car, or su(f). Disjoins regularly from XY, shows 6% nondisjunction from C(1)RM, and causes 9% nondisjunc- tion of + from In(1)dl-49 in In(1)dl-49/+/Dp(1;f)122. # Dp(1;f)134-Dp(1;f)137 origin: X-ray-induced deletion of most of euchromatin from X chromosome marked with y2 or y+. discoverer: Dobzhansky. duplication cytology ref ( genetics ____________________________________________________________________ *Dp(1;f)134 | deletion of most of X 2 y+-br+; not bb+ euchromatin Dp(1;f)135 | deletion of most of X 3 y2-su(s)+ (Voelker); euchromatin bb+ *Dp(1;f)136 / metaphase length 1 y2-pn+; bb+ 1/4 normal X *Dp(1;f)137 metaphase length 1 y2-w+; 1/5 normal X not bb+ ( 1 = Dobzhansky, 1932, Biol. Zentralbl. 52: 493-509; 2 = Dobzhansky, 1935, Z. Indukt. Abstamm. Vererbungsl. 68: 134-62; 3 = Sivertzev-Dobzhansky and Dobzhansky, 1933, Genetics 18: 173-92. | Both sexes viable and wild type except for presence of occipi- tal bristles. / Viability low; shows spread wings and occipital bristles. # Dp(1;f)164 cytology: Dp(1;f)1B;20F (Gersh). Approximate size of chromo- some 4 at metaphase (Yamamoto and Miklos, 1977). origin: X-ray-induced deletion of most of X euchromatin. discoverer: Weltman, 1954. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. Yamamoto and Miklos, 1977, Chromosoma 60: 283-96. genetics: Carries wild-type alleles of y and ac but not sc, su(wa), pn, car, su(f), or bb. Disjoins essentially randomly from XY, shows 36% nondisjunction from C(1)RM, and does not interfere with disjunction of + from In(1)dl-49 in In(1)dl- 49/+/Dp(1;f)164 female. #*Dp(1;f)167 cytology: 3.7-4 times the size of chromosome 4 at metaphase; lacks only the distalmost heterochromatic segment hD (Cooper). origin: X-ray-induced deletion of most of X euchromatin. discoverer: Weltman, 1954. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. genetics: Carries wild-type alleles of y, ac, sc, su(wa), su(f), and bb but not pn or car. Disjoins regularly from XY, shows 3% nondisjunction from C(1)RM, and causes 16% nondis- junction of + from In(1)dl-49 in In(1)dl-49/+/Dp(1;f)167 female. # Dp(1;f)179 origin: X-ray-induced deletion of most of X euchromatin. discoverer: Weltman, 1954. references: Lindsley and Sandler, 1958, Genetics 43: 547-63. genetics: Carries wild-type alleles of y, ac, sc, and su(wa) but not pn, car, su(f), or bb. Disjoins regularly from XY, shows 20% nondisjunction from C(1)RM, and causes 2% nondis- junction of + from In(1)dl-49 in In(1)dl-49/+/Dp(1;f)179 female. # Dp(1;f)749 - Dp(1;f)1518 origin: X ray induced deletion of most of X euchromatin from In(1)sc8. discoverer: Krivshenko and Cooper, 1953. ____________________________________________________________________________ duplication cytology length as ref ( genetics multiple of 4 ____________________________________________________________________________ *Dp(1;f)749 1B12-13;20 3-4 y+-svr+, bb+ Dp(1;f)797 2B4-5;20 2-3 y+-svr+, bb+ Dp(1;f)816 not visible 0.7 2, 3 y+-sc+, bb- Dp(1;f)819 1D3-4;20 2.9 y+-svr+, bb+ Dp(1;f)856 1D3-4;20 3 2, 3, 4, 5 y+-svr+, bb+ Dp(1;f)1144 not visible 1.1 2, 3, 5 y+-ac+, bb- Dp(1;f)1148 2 y+-ac+, bb+ Dp(1;f)1156 2.6 y+-ac+, bb+ Dp(1;f)1158 2.3 y+-ac+, bb+ Dp(1;f)1159 2.7 y+-ac+, bb+ Dp(1;f)1160 3.1 y+-ac+, bb+ Dp(1;f)1162 0.5 2, 3 y+-ac+, bb- Dp(1;f)1170 1.9 y+-ac+, bb- Dp(1;f)1173 3.2-3.6 5, 6 y+-ac+, bb+ Dp(1;f)1185 1.8 y+-ac+, bb+ Dp(1;f)1186 1.6 2, 3 y+-ac+, bb- Dp(1;f)1187 >0.3 | 2, 3, 5 y+-ac+, bb- Dp(1;f)1191 0.7 y+-ac+, bb- Dp(1;f)1193 1 2, 3 y+-ac+, bb- Dp(1;f)1194 3.1 y+-ac+, bb+ Dp(1;f)1201 2.2 y+-ac+, bb+ Dp(1;f)1204 0.9 2, 3 y+-ac+, bb- Dp(1;f)1205 0.7 y+, bb- Dp(1;f)1206 0.5 y+, bb- Dp(1;f)1208 2 y+-ac+, bb+ Dp(1;f)1209 1.9 y+-ac+, bb+ Dp(1;f)1328 / 2A2-3;20 2.1 2, 3 y+-su(wa)+, bb+ *Dp(1;f)1330 2B10-11;20 2.6 y+-svr+, bb+ *Dp(1;f)1331 1E-F;20 1.9 y+-svr+, bb+ Dp(1;f)1337 ` 2B8-9;19-20 1.4 1, 2, 3, 7 y+-hfw+, bb- Dp(1;f)1339 1D-E;20 1.1 2, 3 y+-su(wa)+, bb- *Dp(1;f)1341 2C-D;20 >3 y+-svr+, bb+ Dp(1;f)1342 >3 y+-svr+, bb+ Dp(1;f)1343 1F;20 2.6 2 y+-su(wa)+, bb+ *Dp(1;f)1345 1C;20 1.7 y+-svr+, bb+ Dp(1;f)1346 / 1B12-13;20 2 2, 3, 5 y+-svr+, bb+ Dp(1;f)1479 - 1C;20 2.1 y+-su(s)+, bb+ Dp(1;f)1488 2A;20 2.5 2, 3 y+-su(wa)+, bb+ *Dp(1;f)1489 1D;20 1.8 y+-svr+, bb+ Dp(1;f)1492 / 1B10-12;20 1.9 y+-svr+, bb+ (?) Dp(1;f)1494 1B10-14;20 2.7 y+-svr+, bb+ Dp(1;f)1498 1F;20 3.3 2, 3 y+-su(wa)+, bb+ Dp(1;f)1501 2A;19E4-F1 4.4 y+-svr+, bb+ Dp(1;f)1512 1F;19E4-F1 3.6 y+-svr+, bb+ Dp(1;f)1513 1B10-14;20 >2 y+-svr+, bb+ Dp(1;f)1514 1B12-13;20 1.9 y+-svr+, bb+ Dp(1;f)1518 2A4-B1;20 3.9 y+-svr+, bb+ ( 1 = Belyaeva, Aizenzon, Kiss, Gorelova, Pak, Umbetova, Kra- mers, and Zhimulev, 1982, DIS 58: 184-90; 2 = Grell, 1964, Genetics 50: 151-66; 3 = Grell, 1964, Proc. Nat. Acad. Sci. USA 52: 226-32; 4 = Krider, Yedvobnick, and Levine, 1979, Genetics 92: 879-89; 5 = Parry and Sandler, 1974, Genetics 77: 535-39; 6 = Procunier and Tartof, 1978, Genetics 88: 67-79; 7 = Rayle and Hoar, 1969, DIS 44: 94. | 1300 kb (Karpen and Spradling, 1990, Cell 63: 97-107; Karpen and Spradling, unpublished). Assorts randomly with respect to sex chromosomes. / Lethal with In(1)sc4Lsc8R. ` Variegates for dor, hfw, and BRC in male with no Y chromosome. - Includes su(s)+ according to Voelker. #*Dp(1;f)A1: see *Tp(1;3)A1 #*Dp(1;f)A12: see *Tp(1;2;4)A12 # Dp(1;f)AM cytology: Dp(1;f)1C3-4;8C17-D1;9C1-2;16E1-2; 0.2 length of nor- mal X. new order: 1A - 1C3|9C1 - 8D1|16E3 - 20. origin: X ray induced in In(1)AM = In(1)8C17-D1;16E2-3. references: Bender and Barr, 1971, Nature (London) New Biol. 231: 217-19. Kalisch and Hagele, 1973, Chromosoma 44: 265-83. genetics: Semilethal in males. Does not replicate its DNA in synchrony with homologous region of normal X (Kalisch and Hagele, 1973). #*Dp(1;f)eq: Duplication (1;free) from equational producer origin: X-ray-induced deletion of most of euchromatin from X chromosome carrying eq. discoverer: Schultz, 34k4. genetics: Contains y+ to pn+, inclusive, and bb+. Male fertile but rather inviable; has occipital bristles; eyes rough, wings spread, wing veins thickened. Female has occipital bristles; wings straight edged and coarse textured. Female with two duplications occasionally survives and shows extreme spread wings and rough eyes. # Dp(1;f)GE-1 cytology: Dp(1;f)4C7;20D-F. references: Gelbart, 1971, PhD. Thesis, Univ. of Wisconsin. # Dp(1;f)LJ: Duplication (1;free) La Jolla origin: /-ray-induced deletion of most of the euchromatin from In(1)sc29 = In(1)1B;13A2-5. references: Hardy, Lindsley, Livak, Lewis, Siversten, Joslyn, Edwards, and Bonaccorsi, 1984, Genetics 107: 591-610. duplication cytology new order genetics ______________________________________________________________ Dp(1;f)LJ4 na+ Dp(1;f)LJ8 gv; na+ Dp(1;f)LJ9 ( 1B;12A6-10; 1A-1B|13A2-12A10|20 y+ sc29l(1) 13A2-5;20 g+ Ste+ na+ ( Variegates for g, as in Dp(1;f)LJ9/FM7 and Dp(1;f)LJ9/In(1)dl49, y m2 g4 males (Schalet). Dp(1;f)LJ9/YSX.YL, In(1)EN, Df(1)gl, y, g, f, B males viable and fertile (Schalet). # Dp(1;f)R: Duplication (1;free) from Ring X cytology: Dp(1;f)1A3-4;3A1-2;20A1-F. new order: |1A4 - 3A|20.20F - 20A1|. origin: Spontaneous deletion of most of euchromatin from R(1)2. cytology: Schultz, 35d10. synonym: Dp(1;f)Xc2. references: Ananiev and Gvozdev, 1974, Chromosoma 45: 173-91. Gvozdev, Gerasimova, and Birstein, 1974, Mol. Gen. Genet. 130: 251-60. genetics: Covers y to kz but not bb. Variegation for dor, ac, svr, pn, and kz decreased as Y's are added. Variegation for y insensitive to Y's. #*Dp(1;f)R1-*Dp(1;f)R43 origin: X-ray-induced-deletion of most of euchromatin from R(1)2. discoverer: Pontecorvo. synonym: Del(Xc2)1-Del(Xc2)43. duplication cytology new order ref ( _________________________________________________________________ *Dp(1;f)R1 | 1 *Dp(1;f)R35 1A3-4;17A4-5; |17A5 - 20.20F| 2 19F-20A1 *Dp(1;f)R36 1A3-4;17A4-5; |17A5 - 20.20F| 2 19F-20A1 *Dp(1;f)R37 1A3-4;16F2-3; |16F3 - 20.20F| 2 19F-20A1 *Dp(1;f)R38 1A3-4;1F;20 |1A4 - 1F|20.20F - 20A1| 1 *Dp(1;f)R40 1A3-4;1F4-5;20 |1A4 - 1F4|20.20F - 20A1| 2 *Dp(1;f)R41 1A3-4;1F4-5;20 |1A4 - 1F4|20.20F - 20A1| 2 *Dp(1;f)R42 1A3-4;2A2-3;20 |1A4 - 2A2|20.20F - 20A1| 2 *Dp(1;f)R43 1A3-4;1F4-5;20 |1A4 - 1F4|20.20F - 20A1| 2 ( 1 = Pontecorvo, 1942, DIS 16: 65; 2 = Slizynska, 1942, DIS 16: 67. | Deletion of most of X euchromatin. #*Dp(1;f)R53d cytology: Dp(1;f)1A3-4;1F-2A;20. new order: |1A4 - 1F|20.20F - 20A1|. origin: X-ray-induced deletion of most of euchromatin from R(1)2. discoverer: S. Brown, 1953. synonym: Del(Xc2)53d. references: 1955, DIS 29: 70. Brosseau, 1955, DIS 29: 106. genetics: Contains wild-type alleles of y, ac, sc, and su(s); covers Df(1)260-1. Female tolerates two duplications; male tolerates only one. Fly hemi- or homozygous for y and the duplication shows mosaicism for y. There is probably both variegation for y and loss of the duplication. #*Dp(1;f)RA cytology: Dp(1;f)1A3-4;1F-2A;20 (Slizynska). new order: |1A4 - 1F|20.20F - 21A1|. origin: X-ray-induced deletion of most of euchromatin from R(1)2. discoverer: Pontecorvo. references: 1942, DIS 16: 65. # Dp(1;f)sc7.2 origin: Deletion of majority of euchromatin from In(1)sc7 = In(1)B4-6;5D3-6. references: Garcia-Bellido, 1979, Genetics 91: 491-520. genetics: Carries wild-type alleles of neither cv nor car. #*Dp(1;f)sc260-27: Duplication (1;free) scute cytology: Dp(1;f)1A8-B1;19F. origin: Aneuploid segregant from Tp(1;2)sc260-27/+. # Dp(1;f)scS1Lsc4R (L. Robbins) cytology: Dp(1;f)1-1B3;20F, inferred from genetics. origin: Rex-induced mitotic exchange in In(1)scS1Lsc4R. references: Swanson, 1984, Ph.D. thesis, Michigan State Univ. Swanson, 1987, Genetics 115: 271-76. genetics: One of two mitotic exchange products induced by Rex in In(1)scS1Lsc4R [see also Dp(1;1)scS1Lsc4R]. Material between the two NO regions of In(1)scS1Lsc4R is deleted. Recovered at 1-5% as sterile X/Dp male offspring of Rex/+ or Rex/Rex female by In(1)scS1Lsc4R/Y male crosses. Recovered as X/Dp/Y males at a rate of 10-3 from crosses of Rex/In(1)scS1Lsc4R females to X/Y males. Independently gen- erated Dp(1;f)scS1Lsc4R chromosomes recovered from X/Dp/Y males have different bbRex phenotypes. By analogy with YbbRex, the proximal and distal heterochromatic segments are probably intact and in normal order. # Dp(1;f)scV2 (L. Robbins) cytology: Dp(1;f)1-1B3;20F, inferred from genetics. origin: Rex-induced mitotic exchange in In(1)scV2. references: Swanson, 1987, Genetics 115: 271-76. genetics: A Rex-induced mitotic exchange product of In(1)scV2. Material between the two segments of the divided NO region of the inversion is deleted. Recovered as sterile X/Dp male offspring of Rex-bearing females and In(1)scV2/Y males. By analogy with other Rex-generated chromosomes, the hetero- chromatic constitution should be normal except for bbRex muta- tions induced during the exchange event. #*Dp(1;f)wm3: Duplication (1;free) white-mottled cytology: Dp(1;f)3C-D;19-20; breakpoints inferred from genetic data. origin: X ray induced. discoverer: Muller, 1925. references: 1930, J. Genet. 22: 299-334. genetics: w/Dp(1;f)wm3 male has variegated eyes and is sterile; C(1)RM, w/Dp(1;f)wm3 female has variegated eyes and is fertile. # Dp(1;f)Xc2: see Dp(1;f)R #*Dp(1;f)y-sc: Duplication (1;free) for yellow and scute origin: X-ray-induced deletion of most of X euchromatin. cytology: Oliver, 32k21. references: 1937, DIS 7: 19. phenotype: Carries wild-type alleles of y and sc but not pn. # Dp(1;f)y+ cytology: Small fragment from tip of X. references: Roberts, 1969, Genetics 63: 387-404. genetics: Carries y+. # Dp(1;f)wm51bLwm4R (L. Robbins) cytology: Dp(1;f)1-3C1;20F, inferred from genetics. origin: Rex-induced mitotic exchange in In(1)wm51bLwm4R. references: Swanson, 1987, Genetics 115: 271-76. genetics: One of two mitotic exchange products induced by Rex in In(1)wm51bLwm4R (see also Dp(1;1)wm51bLwm4R). Material between the two NO regions of In(1)wm51bLwm4R is deleted. Recovered at 1-5% as sterile X/Dp male offspring of Rex/+ or Rex/Rex female by In(1)wm51bLwm4R/Y male crosses. By analogy with Dp(1;f)scS1Lsc4R/Y and YbbRex, independent occurrences probably are bbRex with the other heterochromatic segments intact and in normal order. # Dp(1;f)z9: Duplication (1;free) zeste cytology: Dp(1;f)3E7-F1;19-20. origin: X-ray-induced deletion of most of euchromatin from z- bearing X chromosome. discoverer: Gans. references: 1953, Bull. Biol. France Belg., Suppl. 38: 1-90. Jack and Judd, 1979, Proc. Nat. Acad. Sci. USA 76: 1368-72. genetics: Contains z and wild-type alleles of y through dm. Eye color zeste in z w+ males carrying the duplication.